Help debugging an ng-transclude error

I’m trying to hunt down the root of an ng-transclude exception (below). The info in the console doesn’t give any hint about where the problem lies. Does anyone have an idea for how to debug such errors?

Error: [ngTransclude:orphan] Illegal use of ngTransclude directive in the template! No parent directive that requires a transclusion found. Element:
http://errors.angularjs.org/1.4.3/ngTransclude/orphan?p0=<g%20ng-transclude%3D"">
at http://code.ionicframework.com/nightly/js/ionic.bundle.js:8900:12
at ngDirective.link (http://code.ionicframework.com/nightly/js/ionic.bundle.js:36703:35)
at invokeLinkFn (http://code.ionicframework.com/nightly/js/ionic.bundle.js:17482:9)
at nodeLinkFn (http://code.ionicframework.com/nightly/js/ionic.bundle.js:16982:11)
at compositeLinkFn (http://code.ionicframework.com/nightly/js/ionic.bundle.js:16373:13)
at compositeLinkFn (http://code.ionicframework.com/nightly/js/ionic.bundle.js:16377:13)
at publicLinkFn (http://code.ionicframework.com/nightly/js/ionic.bundle.js:16248:30)
at boundTranscludeFn (http://code.ionicframework.com/nightly/js/ionic.bundle.js:16392:16)
at controllersBoundTransclude (http://code.ionicframework.com/nightly/js/ionic.bundle.js:17009:18)
at ngDirective.link (http://code.ionicframework.com/nightly/js/ionic.bundle.js:36710:5)